Unlike compute-rich giants, AppLovin's bootstrapped culture enforces extreme efficiency in its AI infrastructure. Engineers don't have unlimited GPUs, forcing them to optimize code and models for cost and performance. This constraint-driven approach leads to significant cost savings and a lean operational model.

The progression from early neural networks to today's massive models is fundamentally driven by the exponential increase in available computational power, from the initial move to GPUs to today's million-fold increases in training capacity on a single model.

China is gaining an efficiency edge in AI by using "distillation"—training smaller, cheaper models from larger ones. This "train the trainer" approach is much faster and challenges the capital-intensive US strategy, highlighting how inefficient and "bloated" current Western foundational models are.

The perception of LORAs as a lesser fine-tuning method is a marketing problem. Technically, for task-specific customization, they provide massive operational upside at inference time by allowing multiplexing on a single GPU and enabling per-token pricing models, a benefit often overlooked.

The model uses a Mixture-of-Experts (MoE) architecture with over 200 billion parameters, but only activates a "sparse" 10 billion for any given task. This design provides the knowledge base of a massive model while keeping inference speed and cost comparable to much smaller models.

Model architecture decisions directly impact inference performance. AI company Zyphra pre-selects target hardware and then chooses model parameters—such as a hidden dimension with many powers of two—to align with how GPUs split up workloads, maximizing efficiency from day one.

The primary driver for fine-tuning isn't cost but necessity. When applications like real-time voice demand low latency, developers are forced to use smaller models. These models often lack quality for specific tasks, making fine-tuning a necessary step to achieve production-level performance.

The cost to achieve a specific performance benchmark dropped from $60 per million tokens with GPT-3 in 2021 to just $0.06 with Llama 3.2-3b in 2024. This dramatic cost reduction makes sophisticated AI economically viable for a wider range of enterprise applications, shifting the focus to on-premise solutions.

Fine-tuning remains relevant but is not the primary path for most enterprise use cases. It's a specialized tool for situations with unique data unseen by foundation models or when strict cost and throughput requirements for a high-volume task justify the investment. Most should start with RAG.